Yes, I think the Zener diode level shifters may be an important piece to this puzzle. Without the Zeners, i.e. with the simple resistor dividers as level shifters, I'm up against a no-win situation as regards gain. The tubes must be operated near voltage midpoint, i.e., with large cathode resistors. This implies gain of 1. In this configuration, all three resistances, i.e., the plate resistor, the effective resistance of the tube plate-cathode path, and the cathode resistor, are approximately equal. To shift the plate voltage down to the proper grid voltage, requires at least a 1:1 resistor ratio, i.e., gain of 0.5. There seems to be an inevitable no-win voltage-scaling situation, in the absence of Zeners or other non-resistor level shifting. Operating the tubes so that the target voltage at the grids is below midpoint to increase the DC gain above 1, means the plate output voltage must then be scaled down with a more-aggressive resistor ratio, ...